Sony's PlayStation is to challenge Microsoft's Xbox in its one undisputed area of dominance - the online world.
The PlayStation 3 will be "network ready" out of the box when it launches in November and will offer a range of services similar to Xbox Live.
Speaking to developers, he said online games would transform the industry.
Mr MacDonald, vice-president of Sony Computer Entertainment Europe's Worldwide Studios, said: "The PlayStation 3 has been designed so that the network experience is part of the PlayStation experience.
"It is easy to connect, easy to download files."
He said development studios had to re-think their strategies for games releases, arguing that the online space gave them a powerful new relationship with studios.
He said: "But the relationship will only start at the point of sale in the network-centric world. Developers have to change their whole approach."
The classic puzzle game, Super Puzzle Fighter II Turbo, is now playable in Street Fighter 6's Game Centre, along with other new additions.
The Nerd Stash: “Whether you’re here to gloat about your victories or to see what the fuss is about, we present to you the hardest bosses in the Souls series.”